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The D5500 is only slightly less deep than the D5300.
The D5500 retains the stereo microphone.
Nikon has simplified the mode dial, dropping the individual scene program mode options, which took up a lot of space. The adjustment dial is now on top rather than embedded as well.
Though it has the same specifications as the D5300, the D5500 incorporates a touchscreen display; note the Fn option for bringing up the touch control panel.
The interactive panel has been updated to enable touch operation.
The grip is slightly wider and a little more comfortable to hold.
The camera still has the microphone and remote inputs.
Nikon will offer a kit with the collapsible 18-55mm lens as well as the 18-140mm lens.